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  • 1
    Electronic Resource
    Electronic Resource
    Springer
    Experimental brain research 29 (1977), S. 347-366 
    ISSN: 1432-1106
    Keywords: Horizontal Cells ; Cat retina ; Frequency transfer ; Local oscillations ; Flicker responses
    Source: Springer Online Journal Archives 1860-2000
    Topics: Medicine
    Notes: Summary Intracellular responses of horizontal cells in the retina were recorded from the intact eye in situ of anesthetized and paralyzed cats. The responses to large (15×15 °) high modulation (0.6–0.9) photopic (140–200 cd/m2) sinewave light stimuli could be grouped unequivocally into three distinct classes on the basis of the critical flicker frequency (CFF); Hn-units had a CFF of 25–40 Hz, Hm-units of 55–70 Hz and Hw-units of 95–110 Hz. For some stimulus situations small oscillations could be seen in Hm-responses (resonance frequency of about 43 Hz) and Hw-responses (resonance frequency 95 Hz). Hn-responses never showed such an oscillatory tendency. Frequency transfer functions are presented for the three types of H-units and a phenomenological model is discussed that describes these results quantitatively.
    Type of Medium: Electronic Resource
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  • 2
    Electronic Resource
    Electronic Resource
    Springer
    Experimental brain research 37 (1979), S. 65-71 
    ISSN: 1432-1106
    Keywords: Visual stimulator ; Annulus generator ; Orbiting light spots
    Source: Springer Online Journal Archives 1860-2000
    Topics: Medicine
    Notes: Summary When a light spot moves at high angular velocity along a circular path on a screen, an annulus is perceived. This principle is used in a simple annulus generator consisting of a plane mirror that rotates about an axis making an angle α with the normal in the center point of the mirror. Angle α can be varied with a micrometer screw to change the diameter of the annular path (orbit) followed by the light spot that is projected on a screen via the rotating mirror. This results in variable diameter equal energy annuli. The diameter of the light spot can be changed to manipulate the width of the annulus. Finally, at low angular velocities an orbiting light spot is obtained. It is shown in the companion paper (Molenaar and van de Grind, 1979) that this slow orbiting of the light spot allows a precise study of the fine structure of receptive fields.
    Type of Medium: Electronic Resource
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  • 3
    Electronic Resource
    Electronic Resource
    Springer
    Experimental brain research 37 (1979), S. 253-263 
    ISSN: 1432-1106
    Keywords: Horizontal cells ; Cat retina ; Local circuit neuron ; Retinal microcircuits ; Local anisotropy
    Source: Springer Online Journal Archives 1860-2000
    Topics: Medicine
    Notes: Summary Horizontal (H-)cells were recorded intracellularly in the retinae of optically intact feline eyes in vivo. A small light spot orbiting slowly around the receptive field centers was used to quantify the fine structure and diameter of the receptive fields. Receptive field diameters measured in this way were larger than those measured with centered spots of increasing diameter. All H-units studied showed clearly anisotropic receptive field structures. These results are summarized in polar plots representing the local response generating sites with their corresponding “response plus transport” latencies. It is shown that the anisotropic receptive field properties are not incompatible with the approximately homogenous spatial distribution of H-cell somata reported by Wässle and Rieman (1978) for the axonless type of horizontal cell. Finally it is concluded that each H-cell might be involved in many different locally specialized signal processing activities.
    Type of Medium: Electronic Resource
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  • 4
    Electronic Resource
    Electronic Resource
    Springer
    Experimental brain research 29 (1977), S. 367-385 
    ISSN: 1432-1106
    Keywords: Horizontal cells ; Retinal local circuit neurons ; Cat retina ; Flicker responses ; S-Potentials
    Source: Springer Online Journal Archives 1860-2000
    Topics: Medicine
    Notes: Summary In the cat's retina we analyzed the sinewave light flicker response nonlinearity of Hm-type horizontal cells. (For a description of the three types of dynamic responses of cat retinal H-units see Foerster et al., 1977). For equal decreases in the Hm-response amplitude an increase in stimulus frequency had a much stronger linearizing effect than a decrease in stimulus area. Thus the distortion is not simply proportional to response amplitude. Both Hm- and Hn-units had frequency dependent nonlinear area-response functions. The receptive field of Hm-units increased dramatically with stimulus frequency, e.g. from 1 ° at 1 Hz to 8 ° or more at 44 Hz. Intensity transfer data could be described by the function Lb· (ob+Lb)−1 with b ≈ 1 for Hm-units and b ≈ 0.5 for Hn-units. Distortion values predicted from intensity transfer data were too high. It was also found that higher harmonics were attenuated more than the first harmonic at increasing frequencies. Therefore it appeared that some filtering preceded the amplitude compression stage.
    Type of Medium: Electronic Resource
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  • 5
    Electronic Resource
    Electronic Resource
    Springer
    Experimental brain research 91 (1992), S. 135-150 
    ISSN: 1432-1106
    Keywords: Human motion detection ; Visual psychophysics ; Visual distance invariance
    Source: Springer Online Journal Archives 1860-2000
    Topics: Medicine
    Notes: Summary Since visual movement information is often presented in electronic displays or films it is amazing that there is a paucity of research on the influence of viewing distance on motion detection in cinematograms. We report a relatively high degree of detection constancy with changing viewing distance for coherent motion in random-pixel cinematograms. A constant performance irrespective of viewing-distance is called ‘distance-invariance’ and for motion detection it proves to hold reasonably well for a relatively wide range of viewing distances both for foveal and eccentric vision. The limits of this viewing-distance invariance are explored as a function of screen velocity. Detection performance is quantified by a theshold signal-to-noise-ratio (SNR-) value, S, which is determined as a function of velocity for a range of viewing distances from 53 to 13476 mm for foveal vision and from 60 to 1925 mm at 24° eccentricity on the nasal horizontal meridian of the right eye's retina. The data can be explained, at least qualitatively, by a model in which a spatial-resolution stack has a stack of velocity-tuned motion detectors at every resolution layer. Such a ‘stack-of-stacks’ model is in line with proposals for contrast-detection stack-models, but it suggests that the usual hypothesis that motion perception is based on the activity of two separate systems, the short-range and the long-range system, might be superfluous. This two-systems distinction was largely based on the different performance found for moving random dot patterns and moving form-defined stimuli. A moving random pixel array viewed at very close range (e.g. 6 cm) presents the subject with relatively large almost square ‘blobs’, which are less dissimilar from the phi-stimuli used in classic motion perception studies than random dot stimuli at the usual medium to large viewing distances. It leads to maximum displacement threshold (Dm-) values that are not untypical of the ‘long-range’ system, but by gradually increasing the viewing-distance and thus decreasing the pixel-size a continuous change is found from typical long-range to typical short-range values of Dm. The two-systems distinction for motion detection appears to refer to the stimulus rather than to the visual system: The motion-detection system might be forced into a local or a global ‘mode of operation’ by the choice of stimulus.
    Type of Medium: Electronic Resource
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  • 6
    Electronic Resource
    Electronic Resource
    Springer
    Experimental brain research 93 (1993), S. 312-323 
    ISSN: 1432-1106
    Keywords: Plaid motion ; Intersection of constraints ; Texture ; Human
    Source: Springer Online Journal Archives 1860-2000
    Topics: Medicine
    Notes: Abstract Two superimposed gratings, which differ in orientation and move independently, combine into a coherently moving plaid if the component gratings are similar. The effect on the plaid motion percept of the addition of texture to each grating was investigated. The texture disambiguates the motion of each component grating. Under the assumption of recombination of first-stage motion detectors into a second stage, which becomes available to perception, one would expect the perceived motion direction of the plaid to change as a result of texture addition. Subjects perceived the oblique motion direction of textured bars of a single orientation correctly. This occurred for texture details with dimensions down to the resolution limit in the fovea (1 min of arc). Two bar patterns with fine texture (1 min of arc details) which differ in orientation were perceived to cohere into a plaid. The plaid's motion direction, however, was independent of the parallel motion components of the bars. For coarser textures (2 and 4 min of arc details) the bar patterns were perceived to slide past one another. In addition, we found that the plaid motion percept occurred less frequently for longer motion sequences, wider bars and for a combination of the component textures at the intersections which is compatible with partial transparency of the bars. These results do not support the two-stage model of Adelson and Movshon (1982), where only the motion component perpendicular to each grating orientation is encoded and where the perception of the plaid motion results from “recombination” of these perpendicular motion components. The data are more in line with a model where first-stage motion detectors are orientation selective but without the restriction that their preferred direction of motion is perpendicular to their preferred orientation. In the second stage it is proposed that combination occurs across the orientation dimension only. This preserves the direction of motion “labels” at the output of the second stage and allows for representation of transparent as well as coherent plaid motion at this stage of processing.
    Type of Medium: Electronic Resource
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  • 7
    Electronic Resource
    Electronic Resource
    Springer
    Pflügers Archiv 383 (1980), S. 173-179 
    ISSN: 1432-2013
    Keywords: Visual stimulation ; Light stimulus positioning ; Mechanical oscilloscope
    Source: Springer Online Journal Archives 1860-2000
    Topics: Medicine
    Notes: Abstract A “mechanical oscilloscope” is described in which a lightbeam (Z-axis) is projected via a single front aluminized moving mirror. The mirror is mounted in a suspension frame allowing the simultaneous rotation about two mutually perpendicular axes. The linear displacements of two loudspeaker-coils are transformed mechanically into mirror rotations and an electronic position/velocity feedback control system is used to drive the loudspeakers. The bandwidth of the resulting X- and Y-axis deflection systems is of the order of 100–200 Hz. The flexibility of Z-axis control (colour, pattern, intensity, etc.) makes this instrument an attractive component of light stimulation systems for psychophysical or electrophysiological studies of vision.
    Type of Medium: Electronic Resource
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  • 8
    Electronic Resource
    Electronic Resource
    Springer
    Biological cybernetics 8 (1971), S. 85-105 
    ISSN: 1432-0770
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology , Computer Science , Physics
    Notes: Summary Some extensions of the theory of adapting coincidence scaling are presented in the context of neural theory and modelling. Previously the theory of adapting coincidence scaling has been successfully applied to quite a number of specific problems mainly drawn from psychophysical theories of vision: van de Grind et al. (1970a, b); Koenderink et al. (1970a, b). Here emphasis is on neurophysiological problems and after a brief discussion of the “coding” and “component” problems of neural network modelling and a survey of basic coincidence scaling mechanisms a paradigm for neural encoding is treated in some detail. This paradigm (Fig. 6A) is similar to the neuromimes developed and studied by Harmon (1959, 1961) and Küpfmüller and Jenik (1961) for deterministic input signals. On the basis of the introductory discussion of the coding problem it is assumed that the neural code in the peripheral part of the nervous system that we choose as our hunting ground, viz. the retina, is an average event rate code with a Poisson point process as a carrier. Thus the paradigm for neural encoding is studied for such a stochastic input point process. It is then among other things shown that such a simple encoder can generate a wide variety of multimodal interval distributions for certain choices of its parameters. Next we turn to a classic coincidence model of vision and give extremely accurate simulation results to substitute for the lacking analytic solution of the underlying K-fold coincidence problem. A shortcoming of this model is analysed in terms of elementary neural operations and it is shown that the problem of specifying a generalized version of the model ties in with the problem of developing models to explain the quantal signals (bumps) observed on the generator potential during intracellular recordings from the eccentric cell of Limulus. A cybernetic principle for “bump” size adaptation is formulated on the basis of the apparent and possibly significant similarity of this adaptation process with the event rate reduction principle embodied in the so called V R-machine (van de Grind et al., 1970a) which is one of our set of adapting coincidence scalers.
    Type of Medium: Electronic Resource
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  • 9
    Electronic Resource
    Electronic Resource
    Springer
    Biological cybernetics 8 (1971), S. 105-122 
    ISSN: 1432-0770
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology , Computer Science , Physics
    Notes: Summary In the context of the quantum theory of vision scalers, coincidence scalers, adapting coincidence scalers and dead time mechanisms have been used as basic constituents of network models: van de Grind et al. (1970a), Koenderink et al. (1970a, b). The possibilities that these devices offer to construct network models of vision are presently further analysed. First of all a mechanistic analysis is given of the event rate reduction characteristics of “dead time boxes”. Next the interaction of these devices with scalers is discussed in relation with a number of “fluctuation models” of vision proposed in the literature. A critical evaluation of these fluctuation models shows an important defect of most of them, viz. that unrealizable detection criteria are postulated. Our reconsideration of this detection problem then leads to the proposal of some specific realizable detectors. An application of the developed theory of mechanisms (‘machines’) to the explanation of the flash detection characteristics of Limulus concludes the paper. Applications of the presented ideas to neural theory and modelling are treated hi a separate paper (van de Grind et al., 1970b) and for applications of the theory to psychophysically oriented visual modelling studies the reader is referred to Koenderink et al. (1970a, b) and van de Grind et al. (1970a).
    Type of Medium: Electronic Resource
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  • 10
    Electronic Resource
    Electronic Resource
    Springer
    Biological cybernetics 8 (1971), S. 128-144 
    ISSN: 1432-0770
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology , Computer Science , Physics
    Notes: Summary A tentative model of information processing in the human fovea at photopic luminances is described, that handles both luminosity and chromaticity signals. The model consists of a “scaling-ensemble”, a group of sealers with common scaling-factor that provide an effective compression of the dynamic range of the input signals. The scaling factor adapts in such a way that the model conforms to Weber's law for the detection of short, small flashes on a bright background. The dynamics of the adaptation is such that the model effectively computes the logarithmic time derivative of the input signal. The model predicts the outcome of several psychophysical experiments. The predictions include Weber's law, Bloch's law, the apparent brightness of suprathreshold flashes as a function of adaptation level, the influence of spatial inhomogeneities on the perception of flicker, and the transfer function for moving sinusoidal bar patterns for both luminosity and chromaticity modulations. The influence of involuntary eye-movements on the perception of spatial patterns is also discussed. Finally an attempt is made to locate the components of the model in the anatomical structure of the retina. A tentative scheme of neural connections in the human fovea is presented.
    Type of Medium: Electronic Resource
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