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Notes: Abstract.  The predictability of atmospheric responses to global sea surface temperature (SST) anomalies is evaluated using ensemble simulations of two general circulation models (GCMs): the GENESIS version 1.5 (GEN) and the ECMWF cycle 36 (ECM). The integrations incorporate observed SST variations but start from different initial land and atmospheric states. Five GEN 1980–1992 and six ECM 1980–1988 realizations are compared with observations to distinguish predictable SST forced climate signals from internal variability. To facilitate the study, correlation analysis and significance evaluation techniques are developed on the basis of time series permutations. It is found that the annual mean global area with realistic signals is variable dependent and ranges from 3 to 20% in GEN and 6 to 28% in ECM. More than 95% of these signal areas occur between 35 °S–35 °N. Due to the existence of model biases, robust responses, which are independent of initial condition, are identified over broader areas. Both GCMs demonstrate that the sensitivity to initial conditions decreases and the predictability of SST forced responses increases, in order, from 850 hPa zonal wind, outgoing longwave radiation, 200 hPa zonal wind, sea-level pressure to 500 hPa height. The predictable signals are concentrated in the tropical and subtropical Pacific Ocean and are identified with typical El Niño/ Southern Oscillation phenomena that occur in response to SST and diabatic heating anomalies over the equatorial central Pacific. ECM is less sensitive to initial conditions and better predicts SST forced climate changes. This results from (1) a more realistic basic climatology, especially of the upper-level wind circulation, that produces more realistic interactions between the mean flow, stationary waves and tropical forcing; (2) a more vigorous hydrologic cycle that amplifies the tropical forcing signals, which can exceed internal variability and be more efficiently transported from the forcing region. Differences between the models and observations are identified. For GEN during El Niño, the convection does not carry energy to a sufficiently high altitude, while the spread of the tropospheric warming along the equator is slower and the anomaly magnitude smaller than observed. This impacts model ability to simulate realistic responses over Eurasia and the Indian Ocean. Similar biases exist in the ECM responses. In addition, the relationships between upper and lower tropospheric wind responses to SST forcing are not well reproduced by either model. The identification of these model biases leads to the conclusion that improvements in convective heat and momentum transport parametrizations and basic climate simulations could substantially increase predictive skill.