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Notes: [Auszug] Table 1 Intake and digestibility of diets DMI DMI/ DMD MRT kgW0.75 (g) (g) (%) (h) January Red deer Agrostis/Festuca 1,199 49.1 40.1 27.4 Heather 847 35.0 45.3 37.7 Sheep Agrostisl Festuca 432 24.1 48.8 56.1 Heather* 327 17.7 ...

Notes: The effects on herbage intake and ingestive behaviour by ewes and weaned lambs of grazing aftermath and previously continuously grazed perennial ryegrass-dominant swards at two different sward heights (4 and 8 cm) in the autumn were studied. The experiment had a factorial design, was replicated twice and was conducted from mid-August to early November. There were six ewes and six weaned lambs per treatment plot and measurements were made in three periods. The effects of previous treatment of swards on herbage intake by ewes and weaned lambs were greatest in August, with herbage intakes being significantly lower on the aftermath swards. Differences disappeared by October. The lowest herbage intakes were obtained on the aftermath sward at the lowest sward height, with ewes being more affected under those conditions than lambs. Grazing time and biting rate increased with a reduction in sward height and were higher on aftermath swards. However, these increases did not compensate for reductions in estimated bite size on the aftermath swards. It was concluded that the effects of the sward management treatments in the summer on tissue turnover of the sward and herbage intake in the autumn were considerable in the early part of the autumn but had largely disappeared by the end of the autumn period.

Notes: A series of twenty-four swards containing different proportions of white clover (0·20-0·25) and perennial ryegrass were created by using different seed mixtures, herbicide applications and previous cutting Frequencies. These swards were used to study the diet of oesophageally-fistulated wether sheep which grazed the various swards for a 30-min period after 1, 2 and 3 weeks of regrowth.The proportion of white clover in the diet was generally greater than that in the sward. Fifty-seven percent of the variation in the proportion of white clover in the diet could be attributed to the proportion of white clover in the sward. White clover and perennial ryegrass leaf and stem were grazed to the same height and the proportion of white clover in the grazed horizon of the sward explained 83% of the variation in the proportion of white clover in the diet. The proportion of white clover in the diet was greater than the proportion in the grazed horizon of the sward in week 3 of regrowth, but not in weeks 1 and 2, and greater when the proportion of white clover in the grazed horizon was lower than 0·20. Both these observations were interpreted as indicating selection for white clover by the sheep within the grazed horizon.There was a positive and linear relationship between the depth of the grazed horizon and sward height which, together with the relationship between the proportion of white clover in the grazed horizon and in the diet, would allow the prediction of the proportion of white clover of the diet from the height and the white clover content of the grazed horizon of the sward.

Notes: A description is given of equipment designed to be fitted to either sheep or cattle, and to record automatically information on the jaw movements associated with both grazing and ruminating, movements of the head in plucking mouthfuls of herbage, and the time spent in grazing. The equipment has been successfully tested against alternative visual and mechanical methods of measuring grazing behaviour.The use of modified equipment in detailed studies has shown (a) that the maximum acceleration of the head in a longitudinal plane when plucking mouthfuls of herbage was twice as great in sheep as in cattle and in the sheep, but not in the cow, increased as sward height decreased; and (b) that the ratio of jaw movements to head movements was always greater than unity in both sheep and cattle, and was greater on tall than on short swards in two out of three comparisons. The rate of biting was significantly lower on tall than on short swards.

Notes: The effects of sheep grazing on the development and production of heather swards were studied over a period of 3 years in an experiment in which sheep numbers were adjusted to remove 0, 40% or 80% by weight of the current season's shoots in summer and in autumn.The number of shoots grazed increased with the level of grazing but not in direct proportion. The length of shoot grazed was greater at the 80% level of grazing. Heavy autumn grazing was associated with frequent uprooting and breaking off of shoots and with a high incidence of shoots grazed down to the previous season's wood. The siting of new season's shoots was influenced by the amount of shoot length grazed in the previous season. Grazing behaviour of sheep within a plot was affected by variation in heather cover with the sheep showing a tendency to graze near bare areas.After 3 years of grazing, standing crop was reduced by both levels of grazing whether in summer or autumn. New shoot production was unaffected on plots which had had the 40% level of shoot removal but was reduced by half on plots which had had the 80% level of shoot removal whether in summer or autumn.A clipping experiment was also carried out where 0, 40% or 80% of the length of current shoots was removed in May, early July and Septeinber. Standing crop was unaffected by May clipping but was reduced on plants with a history of 80% shoot removal in July or September. New shoot production was only reduced on clipped plants which had been subjected to 80% removal in September.The role of overwintering green shoots and carbohydrate reserves in the support of new shoot growth was considered.The implications of the results in relation to heather management are discussed.

Notes: Methods of predicting the in vivo dry matter digestibility (DMD) of heather were examined in studies on ten samples of heather which had been fed to sheep in standard digestibility trials. DMD was predicted with similar accuracy from estimates of in vitro dry matter digestibility (in vitro DMD) and from the concentration of cell-wall constituents. In vitro DMD was found to be between 10 and 18 percentage units lower than DMD. Freeze drying the samples was the only modification to the technique which both increased in vitro DMD and the accuracy of the prediction of DMD. Other modifications attempted were the use of different liquor sources, changes in first-stage incubation time and dilution rate, and nitrogen supplementation of the inocula.The concentration of nitrogen or undigested dietary nitrogen in the faeces could not be used to predict the digestibility of heather with precision. It was concluded that the DMD of a heather diet could be satisfactorily predicted from the in vitro DMD of samples collected from sheep fistulated at the oesophagus.

Notes: The effects of sheep grazing on the development of and production from heather swards and on intake and diet selection by grazing Scottish Blackface wether sheep were studied in an experiment in which sheep numbers were adjusted to remove 0, 40% or 80% by weight of current season's shoots in summer and in autumn for a period of 5 years. All combinations of season and level of grazing were provided. The plots were divided after 2 years and grazing was continued for a further 3 years on one subplot but discontinued on the other.Sward productivity during the 5 years of grazing was unaffected at the 40% level of shoot removal but at the 80% level was reduced by up to 66%.After a season's uninterrupted growth at the end of the experiment heather cover and herbage mass were lower and current season's shoots as a proportion of total mass were higher on those treatments which received the greatest severity of grazing. Weight of current season's shoots was unaffected on treatments which had received the 40% level of shoot removal but was reduced by 40% and 50% on treatments which had received the 80% level in summer and autumn respectively. On the rested subplots sward recovery was such that no treatment effects remained after 3 years except with respect to 80% shoot removal in autumn where herbage mass was reduced compared with other treatments.Intakes of digestible organic matter per sheep were higher at the 80% than the 40% level of grazing in the fifth year of the experiment. This was ascribed to the ingestion of new shoot growth from the twig bases on the 80% level of removal treatments. Intake and digestibility were higher in the summer on those treatments which had received the 80% level of removal in previous autumns. Over the 5 years of the experiment there was a small decline in intake and digestibility values, with the decline being greatest in the summer and at low levels of grazing.Floristic changes differed on the rested and grazed areas. On the rested subplots in the fourth year after grazing stopped, heather height was lower and cover by grasses, sedges and herbs was greater on treatments which previously had received the highest levels of utilization. On bare areas the rate of recovery of heather was rapid where shoots as well as seedlings contributed to recovery growth. On the grazed subplots, with the exception of the low-growing ruderal Rumex acetosella, cover due to grasses, sedges and herbs remained low as species other than heather were selectively grazed.Relationships among stocking rates, grazing behaviour and heather utilization are discussed and guidelines for the recognition of overgrazing are outlined.

Notes: The effect of level of removal of current season's shoots of heather (0,40 and 80% of the dry matter of the shoots) in the summer and autumn on the intake and diet selection by grazing Scottish Blackface wether sheep was examined in an experiment conducted over two years in which all combinations of level and season of grazing were provided. The treatment plots were grazed for 5 weeks at any one time and measurements made of intake and diet selection in weeks 2 and 5.In both the summer and autumn grazing periods level of removal in week 2 had little effect on the quality of the diet selected. In week 5, digestibility of organic matter was higher for the 40% than the 80% level of removal in the summer (0·543 vs 0·508) and particularly in the autumn (0·503 vs 0·449). Digestibility of the diet selected was higher in week 2 than in week 5 in both the summer (0·555 vs 0·525) and in the autumn (0·511 vs 0·476). Mechanisms of diet selection are discussed.In the summer grazing period only small differences existed in intake of organic matter between levels of removal but in the autumn intakes were higher at the 40% than the 80% level of removal. In a clipping experiment in which 0, 40 or 80% of the length of current season's shoots was removed in May, July and September in 3 consecutive years the nitrogen content of clippings in May and July was higher and the cell wall constituent lower at the 40% level in comparison to the 80% level of removal. Higher levels of removal at the previous clipping increased the nitrogen content of the clippings in May, July and September and reduced the cell wall constituent content in September.

Notes: Abstract The objective of the study was to describe the annual and seasonal production of live biomass of six plant species or communities of the uplands of the UK –Calluna vulgaris, Vaccinium myrtillus, Nardus stricta, Molinia caerulea, Eriophorum vaginatum and Agrostis–Festuca grassland. The species or communities are important for the management of the uplands to meet both biodiversity and agriculture objectives. The annual and seasonal production of live biomass was determined for sites within six regions of England and Wales in 1995, 1996 and 1997. The six regions were North-east England, North-west England, North Pennines, South Pennines, South-west England and Wales. In each region, the Institute of Terrestrial Ecology's Land Classification Scheme was used to select three or four 2 km × 2 km squares that contained a mosaic of C. vulgaris and upland grassland plant species. Uniform plots of 1 ha were selected from each 2 km × 2 km square, within which three measurements were made on each sampling occasion. Biomass of each species or community was measured within 1·0 × 0·5-m quadrats using the same standard procedures. Measurements of annual production of current season's shoots of the pioneer, building, mature and degenerate phases of C. vulgaris and of V. myrtillus were made in October, and of the annual production of live biomass of N. stricta, M. caerulea and E. vaginatum in August. A simulated grazing regime was imposed for the Agrostis–Festuca grassland with cuts being made to a height of 4 cm at 8-week intervals. The annual production of live biomass produced was estimated as the sum of four harvests taken during the growing season. Seasonal measurements of the amount of live biomass of all the species were also made in April, June, August and October. For C. vulgaris, there were significant (P = 0·05) differences between the pioneer, building, mature and degenerate phases in the annual production of current season's shoots, with greater amounts being produced by the phases of greater age and with higher biomass. There were significant year (P 〈 0·05) and region (P 〈 0·05) effects, with the lowest production in 1995 and in the South-west of England. The values were higher than those observed in previous studies in Scotland, and these were interpreted as being associated with a longer growing season rather than higher summer temperatures. There was no difference between years and regions in the annual production of V. myrtillus. For the grass species, there were small differences in annual production of live biomass between years and regions. However, the annual production of live biomass of N. stricta was significantly (P 〈 0·01) lower in 1995 than in 1996 and 1997 and significantly (P 〈 0·001) lower in the South Pennines region than elsewhere. The effect of a range of weather and other environmental variables on the annual production of live biomass of each species of grass was explored, but they provided only a limited insight into the reasons for the observed differences and were found to be of limited value as predictors.